Kleinschmidtimyia Spencer, 1986
The seven species placed in this genus were initially described as Melanagromyza and Ophiomyia (Kleinschmidt, 1961, Spencer, 1986). The larval and imaginal morphology indeed indicates a close relationship with Melanagromyza. Later (Spencer, 1973) transferred some of them to Tropicomyia and described subsequently some new species. He did so because of the species' unusual feeding habit. As other Tropicomyia they are leaf miners feeding mainly on the epidermal layer. However, they lack the unusual serrated mandibles typical for Tropicomyia and have normal alternating mouthhooks instead.
Because of the unusual life history, the missing commonalties with Tropicomyia and, mainly, the distinctive genitalia a new genus was erected (Spencer, 1986).
Immature stages
Both larvae and adults of Kleinschmidtimyia are typical members of the Ophiomyia-group (Tribe Ophiomyiini), they have the usual gonites, subbasal vesica of distiphallus and the black species have dark halteres and two rows of two dorsocentral bristles on the mesonotum. Puparia always remain in the larval mine and have as far as we know the additional longitudinal suture on the upper puparial cap. The cephalopharyngeal skeleton including the arrangement of the mandibles is as in the whole group of the Ophiomyiini.
Male genitalia
The male genitalia show some apparently specific characters of Kleinschmidtimyia: The hypandrial apodeme is always developed and extraordinary long (Kl pisi terminalia.pct); the distiphallus has the same structure as in the other genera but with elongated terminal tubules (Kl pisi aedeagus.pct). The basiphallus resembles the one in Melanagromyza and Tropicomyia in the apparent symmetry.
Distribution
The genus is known from Australia and New Guinea only.